A direct evidence of the effect of egg yolk in oil paint, or lack thereof, can be seen in Leonardo da Vinci’s “Madonna of the Carnation,” one of the paintings observed during the study.

Leonardo da Vinci paints The Virgin on the Rocks – the most complex Madonna image of the entire Renaissance. But, after a disagreement over money, he withheld the painting.

Madonna of the Yarnwinder (The Buccleuch Madonna) (c. 1499–1508) Leonardo da Vinci, The Madonna of the Yarnwinder (ca. 1501). Photo by Fine Art Images/Heritage Images/Getty Images.

LEONARDO DA VINCI looks at how the artist influenced ... The Virgin, infant Jesus, John the Baptist and an angel also called "The Madonna of the Rocks" or "Virgin of the Rocks." ...

Leonardo da Vinci; The Madonna and Child with the Infant Saint John the Baptist; 71.8 x 50.5 cm. (28.3 x 19.9 in.) close. Medium oil and gold on panel Size 71.8 x 50.5 cm. (28.3 x 19.9 in.) Sale ...

His arm is proudly wrapped around a painting - Leonardo da Vinci's The Madonna of the Yarnwinder. The Missing Madonna podcast is available on BBC Sounds. Related topics. Leonardo da Vinci; ...